Tiny Brains, Giant Impact: Uncovering the Keystone Neurons of LLM with Just a Few Prompts

Large language models (LLMs) display strong comprehensive abilities, yet the internal mechanisms that support these behaviors remain insufficiently understood. In this work, we show that across a wide range of open-weight Transformers, a subset of neurons remains consistently highly activated during inference across tasks of multiple capability dimensions. By probing along the cross-task activation strength, an extremely sparse subset is isolated, whose removal causes a collapse in model behavior, which we term keystone neurons. Our analysis reveals that keystone neurons are a stable and intrinsic neuron subset of the model that is largely established during pretraining. The parameters associated with these neurons are tightly calibrated during the training process, and their precise values are critical for the capabilities of the model. Building on these insights, we propose a supervised fine-tuning approach that updates only keystone neurons, achieving task gains comparable to or even better than full-parameter fine-tuning while better preserving performance in other capability dimensions, despite modifying a much smaller number of parameters.

Large language models can perform a wide range of tasks, yet the internal components that support these abilities remain poorly understood. We identify a tiny set of neurons that is consistently active across diverse prompts in many open-weight models. Removing these neurons causes broad capability collapse, which reveals their central role in model behavior. These neurons are stable across prompt choices and appear to emerge mainly during pretraining. We further show that updating only their associated weights enables efficient model adaptation with far fewer parameter changes.